- Z-Mount Lens/FX Format
- Aperture Range: f/4.5 to f/32
- SR, Super ED, and ED Elements
- Nano Crystal Coat
- Stepping Motor AF System
- Vibration Reduction Image Stabilization
- Customizable Fn and Memory Set Buttons
- Weather-Sealed Design, Fluorine Coating
- Rotating Tripod Collar
- Rounded 9-Blade Diaphragm

Distinct among super-telephoto lenses, this 400mm f/4.5 is especially sleek and lightweight, making it an ideal companion for travel or other shooting scenarios where handheld shooting is preferred. Sophisticated optical design includes a series of specialized elements that promote a lighter, more compact overall design. Modest f/4.5 maximum aperture offers a practical balance between size, speed, and weight. With a size more akin to a 70-200mm f/2.8, this lens measures just 9.2" long and weighs 2.7 lb. Compared to the Z 400mm f/2.8 lens, this f/4.5 version weighs 3.8 lb less and measures 6" shorter in length.

Helping to virtually eliminate chromatic aberrations, color fringing, and comatic aberrations, this lens features one ED (extra-low dispersion) glass element and two Super ED elements, along with one SR (short-wavelength refractive) element. The SR element features a specialized high-dispersion design that refracts wavelengths shorter than blue; this helps achieve truly accurate colour rendering and also aids in making the lens design more compact. Nano Crystal Coat has been applied for well-rounded anti-glare protection at all angles, helping to suppress ghosting, reflections, and flare for higher contrast and more accurate color response when working in strong lighting and backlit conditions.

Advanced autofocus system uses stepping motors to deliver quick and accurate focusing performance, along with smooth and quiet AF that is well-suited to both stills and video applications.

- Nikon NIKKOR Z 400mm f/4.5 VR S Lens
- Nikon LC-95B 95mm Snap-On Front Lens Cap
- Nikon LF-N1 Rear Lens Cap
- Nikon HB-105 Lens Hood
- Nikon CL-C5 Lens Case
- Tripod Foot